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CPIE Notebook Project - Grasses of Hawai‘i and Guam Grass Key – Page 16

  Open Paniculate Inflorescences with long awns
[PANICULATE INFLORESCENCES CONTINUED]

     
101a (To last page83) A single, fertile floret above each glume pair. Spikelets with one or two awns .. . [102]
101b

Multiple, fertile florets above each glume pair. Spikelets with more than two awns .. .

[111]
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102a (101)

Inflorescence an erect, panicle of paired spikelets, each greater than 2.5 mm long .. .

[103]
102b Panicles dense, spikelets narrow, up to 2.5 mm long. Medium-size perennial grass with hairy, sticky leaves (particularly the sheaths). Strong-smelling (like molasses). Inflorescence varies from very narrow to narrow ovoid, purple-tinged. { Widely distributed in dry to mesic sites above 400 ft elevation. Molasses grass [HAW - NAT] [FAC]; Molasses grass [GUM - ?NAT] [FAC]
    Melinus minutiflora P. Beauv.
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103a (102) Spikelets 2 to 6 mm long excluding awn, paired: sessile spikelet perfect, awned; pedicellate spikelet not awned .. .

[104]
103b

Spikelets 9 mm (>5/16 in) or more long not including awn(s), which exceed length of spikelet .. .

[107]
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104a (103) Inflorescence of one to many racemes, these typically paired or digitally arranged on erect branchlets; spikelets paired, more or less similar, but only sessile one with long awn. Medium size grasses. Dichanthium spp. .. .

[106]
104b Sorghum spikelets Paired spikelets disimilar, stalked spikelet (two at branch end) narrow, sterile or male only; sessile spikelet with a twisted and bent awn. Leaves wide, with conspicuous midrib. blade and sheath glabrous, except densely pubescent at throat; ligule membranous, dark, conspicuously longest at middle, fringed with short tight hairs.{ Inflorescence varies from crowded to somewhat open, narrow ovoid to spindle-shaped, or drooping. Large grasses. Sorghum spp. .. .


Figure 16A. Photo drawing of (seven) Sorghum spikelets. Note terminal sessile spikelet is associated with two pedicellate spikelets, the others with one each.
[105]
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105a (104) Raceme somewhat fragile. Grain conspicuous, shiny white at maturity. { Large rhizomatous, perennial (Fig. 16A). Johnson grass [HAW - NAT] [FACU]; Johnson grass [GUM - NAT]
    Sorghum halepense (L.) Pers.

105b Raceme tough, tardily disarticulating. Grain large, spreading the glumes (some black), colored reddish brown at maturity. { Large annual. [HAW - NAT] [FACU]; broomcorn [GUM - NAT]
    Sorghum bicolor (L.) Moench
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107a (103) Large perennial with dense, open, but typically drooping inflorescence. Apparent spikelets on mostly stiff branches arising from the leaf sheaths. Actual spikelets are mostly much reduced and number 3-6 above a 3-3.5 cm (>1 in) spathe. Two spikelets with long awns in each complex raceme, although often missing. Conspicuous is a black, rufous pubescent, truncated glume surrounding each spikelet. { Naturalized in dry to mesic pastures and other disturbed sites to 2500-ft elevation. HDA noxious weed. Lyon's grass [HAW - NAT]
    Themeda villosa (Poir.) A. Camus
107b Spikelets single on narrow pedicels branching off the rachis. Medium to large grass with open and not particularly dense inflorescence. Two awns per spikelet. Avena spp. .. . [108]
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108a (107) Rachilla and lemma glabrous or sparesely pubescent. Long awns straight or weakly bent. Florets not deciduous .. . [109]
108b

Rachilla and lemma hairy. Long awns typically bent. Florets readily deciduous, disarticulating above glumes and between floret pair .. .

[110]
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109a (108) Lemma glabrous at base. Long awns typically straight. Tip of lemma entire or slightly bilobed, without awns. Florets not deciduous, persisting. { Medium to large annual grass, cultivated as forage, but naturalized in disturbed places and pastures. Common oat [HAW - ORN,NAT] [FAC] [GUM - ORN]
    Avena sativa L.
109b Lemma glabrous but rachilla of second floret ciliolate on upper edges. Long awns slightly bent. Tip of lemma with two short awns. { Medium to large annual grass, cultivated as forage on Kaua‘i (coll. 2013). Hybrid oat. [HAW - ORN] [FAC]
    Avena sativa hybrid or cross?
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110a (108) Tip of lemma with two short awns. Lemma hairs white. Naturalized on O‘ahu, Maui, and Hawai‘i. [HAW - NAT]
    Avena barbata Pott ex Link
110b Tip of lemma toothed, but without bristles or awns. Lemma hairs yellow to brown. Wild oat [HAW - NAT]
    Avena fatua L.
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111a (101) Spikelets tapering towards the tip. Lemmas up to 0.5 in (1 cm) long and 0.1 in (2.5) mm wide in side view. Lower glume 3-7 veined. Awns straight, about 1/4-in (4-9 mm) long. { found in dry areas above 1,000 ft. [HAW - NAT] [UPL]
    Bromus hordeaceus L.
111b Spikelets gaping somewhat at the tip. Lemmas 0.5 to 1.25 in (1 to 3 cm) long and less than 0.1 in (1-1.5 mm) across in side view. Lower glume with a single vein [112]
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112a (111) Bromus tectorum spikelets Panicle open and drooping. Lemmas 0.5 to 0.8 in (1-2 cm) long, membranous and bifid at the tip, supporting a scabrous awn, straight and 0.5 to 0.7 in (1-1.7 cm) long. { Adventive on East Maui. Cheatgrass [HAW - NAT]
        Bromus tectorum L.

Figure 16B. Inflorescence of B. tectorum.
112b

Spikelets larger, Lemmas 2-3 cm long, bifid at the tip, supporting a scabrous awn 1 to 2 in (30-60 mm) long. { Common in pastures, subalpine forest and shrubland, and disturbed areas. Ripgut brome [HAW - NAT]

    Bromus diandrus Roth


Synonyms for species described on page 16:

    Bromus mollis L. (= B. hordeaceus)
    Bromus rigidus Roth (= B. diandrus)
    Bromus willdenowii Kunth (= B. catharticus)

Species from Hawai‘i not included in the key:

    Bromus carinatus Hook. & Am.— Faccenda, 2022.
    Bromus japonicus Thunb. — Faccenda, 2022.
    Bromus madritensis L. — Faccenda, 2022.
    Bromus rubens L. (= B. madritensis in Hawai‘i)— Faccenda, 2022; QUESTIONABLE – Faccenda 2024b
    Bromus secalinus L.— Faccenda, 2022; QUESTIONABLE – Faccenda 2024b
    Bromus sterilis L. — Faccenda, 2022.
    Sorghum arundinaceum (Desv.) Stapf (= Sorghum xdrummundii)
    Sorghum xdrummundii (Nees ex Steud.) Millsp.

Species from the Marianas not included in the key:

    Sorghum xdrummundii (Nees ex Steud.) Millsp.
    Sorghum nitidum (Vahl) Pers.
    Sorghum propinquum (Kunth) Hitchc.
    Sorghum sudanense (Piper) Stapf ex Prain (= S. xdrummundii)


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