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40a |
(26 & 38) |
Inflorescence an umbrella like arrangement of (1-)3-9 spikes. Each spike with two rows of downward-pointing bristled spikelets appearing like teeth on a comb. Rachis usually extending as a conspicuous point beyond last spikelets. Ligule a fimbriate membrane. { Widespread although typically not abundant where found; small running grass (Fig. 12B). Beach wiregrass [HAW - NAT] crowfoot grass [GUM - NAT] Dactyloctenium aegyptium (L.) Willd.
Figure 12B. Beach wiregrass (Dactyloctenium aegyptium) inflorescence of spikes arranged with the spikelets pointing downward and comb-lkike due to bristles.
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40b
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Inflorescence of spikes or racemes, with or without bristles, but not comb-like in appearance, the spikelets tending to lay flat against the rachis, the tip of which is hidden by uppermost spikelets .. .
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[41]
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41a
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(40) |
Spikelets with awns (bristle-like narrow appendage arising near the upper end of one or more bracts of the spikelet) and at least as long as the supporting bract. (Note: true bristles are stiff hairs arising from the base of the spikelet, not from within the spikelet).
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[48]
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41b
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Spikelets without awns or awns very short (less than length of the bracts surrounding the floret) and not conspicuous even with low-power magnification
.. .
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[42]
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42a
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(41) |
Inflorescence of 3 to 20 coarse spikes or racemes. Small to medium spreading or clumping grass .. .
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[43] |
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42b
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Inflorescence of 3 to 9 delicate, narrow spikes or racemes. Small grass spreading by stolons or rhizomes or both
.. .
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[45] |
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43a
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(42) |
Small to medium clumping grass. Spikelets 3+ mm long. Ligule a short (0.5 mm), erose or ciliate membrane .. .
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[44] |
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43b
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Coarse, medium, spreading grass. Spikelets 3 mm or less in length. Ligule a dense fringe of hairs
.. .
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[47] |
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44a
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(43) |
Culms expanded at base, conspicuously flattened and often whitish. Ligule a short, erose membrane. Inflorescence digitate, but often with one raceme located on culm an inch or two below the 2-6 clustered racemes. Spikelets nearly 1/4 inch (5-7 mm) long, appearing coarsely toothed, but not awned, compressed, and greatly overhanging the rachis. All bracteoles glabrous with membranous (scarious) margins. { Found typically as a weed growing prostrate in mowed lawns, but upright to 1 or even 2 m where not so disturbed. Wiregrass [HAW - NAT] [FACU], umog, goosegrass [GUM - NAT]Eleusine indica (L.) Gaertn.
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44b
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Inflorescence of (5-)7-20 spikes. Spikelets with several short awns, the longest about as long again as a spikelet, the shortest very short and on the upper glume; some bractioles ciliate. { Medium perennial grass of open, disturbed areas, verges, and pastures (Fig. 12A). (See also couplet [48a]) Rhodes grass [HAW - NAT] [FACU] [GUM - NAT]
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45a
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(42) |
Spikelets narrowly ovate with one or more awns. Leaf arrangement distinctly alternate. Some Chloris spp.
.. .
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[50] |
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45b
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Spikelets ovate, broadly or not, but without awns. Leaf arrangement alternate or distichous (two-ranked; alternating blades coming off culm in two vertical rows on opposite sides of the axis) .. .
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[46] |
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46a
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(45) |
Plants 5-40 cm tall (to 16 in). Inflorescence of usually 3-6, stiff, thin spikes in a single whorl. Spikelets broadly ovate, compressed, without awns, falling without glumes (see Fig, 12D, below) and under 1/16 in (3.5-4.5 mm) long. Leaf arrangement distinctly distichous (Fig. 12C, left), especially on short, vegetative stalks. { Small (but quite variable in size), perennial grass spreading by stolons and rhizomes, widely used as a lawn grass. Bermuda grass, mānienie [HAW - NAT] [FACU]; Bermuda grass, grama [GUM - NAT] [FAC] Cynodon dactylon (L.) Pers.
If plant is > 20 cm tall and number of branches greater than 5 in one or more whorls, consider also couplet [47]. In a planted lawn situation, see Page 21.
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46b
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Spikelets narrowly ovate, pediceled, pedicels unequal, spikelets falling at maturity with glumes. Leaf arrangement distinctly alternate. Some Digitaria spp. .. .
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[36] |
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47a
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(43) |
Culms soft, not woody. Inflorescence of 3-13 spikes in one whorl (sometimes two). Spikelets single, on short pedicels borne alternately on either side of a flatened, curved rachis. African Bermuda grass [HAW - NAT]Cynodon nlemfuensis Vanderyst
Although rare, these two species may hybridize in Hawai‘i (Faccenda, et al., 2024)
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47b
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Culms stiff, "woody". Inflorescence of 5-17 spikes in one to five whorls. Giant star grass [HAW - NAT] Cynodon aethiopicus Clayton & J.R. Harlan
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48a
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[41] |
Spikelets under 6 mm (1/4 in) and crowded or not on the rachis of the raceme. Unequal glumes remaining on the rachis after the spikelet falls away .. .
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[49] |
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48b
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Spikelets large (to 13 mm or 1/2 in) and so dense on a rachis at maturity that the rachis itself is obscured. Racemes may curve inward at tips. Spikelets not disarticulating at maturity. { Resembles the very common E. eleusine but much more robust; a large grass; seeds are exposed at maturity and edible; plant is grown as a crop in parts of Africa and Asia. ragi, ragimillet [GUM - NAT] Eleusine coracana (L.) Gaertn.
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49a
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[48] |
Spikelets strongly laterally compressed, linear or lanceolate in outline on short pedicels crowded in two rows on rachis. Lower lemma with a bent awn, twisted at the base. Glumes keeled. Ligule a fringed membrane .. .
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[50] |
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49b
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Spikelets rounded or linear, not strongly compressed, on short or no pedicels. Glumes not keeled. Ligule a membrane or dense row of cilia. Chloris spp. .. .
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[45] |
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50a
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[49] |
Medium-size, grey-green grass; plant densely pubescent. Racemes 3 to 6. Leaves slender. Rachis trigonous. Spikelets 3.3-4.8 mm long, inserted into alternating notches in the rachis. Tuft of short hairs at base of each spikelet. Apex of lower lemma bifid. { Abundant perennial grass of grasslands and savanna on Guam. [GUM - END] Dimeria chloridiformis (Gaud.) K. Schum. & Lauterbach
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50b
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Small to medium-size grass; plant glabrous or sparsely pubescent. Racemes paired, short. Leaves short. Rachis flat, glabrous on the margins. Spikelets 0.8-3 mm long. Lower lemma awn 7-9 mm long. Upper glume deeply forked. { Annual. Mostly found in ditches and fallow rice fields. [GUM - IND]Dimeria ornithopoda Trinius
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